Bennett, D.C. and Lamoreux, M.L. (2003). The color loci of mice–a genetic century. Pigment Cell Research, 16: 333-344. doi.org/10.1034/j.1600-0749.2003.00067.x.
Bultman, S. J., Michaud, E. J. and Woychik, R. P. (1992). Molecular characterization of the mouse agouti locus, Cell, 71: 1195–1204. doi.org/10.1016/s0092-8674(05)80067-4.
Cieslak, M., Reissmann, M., Hofreiter, M., and Ludwig, A. (2011). Colours of domestication, Biological Reviews, 86: 885–899. doi.org/10.1111/j.1469-185X.2011.00177.x.
Costin, G. E. and Hearing, V. J. (2007). Human skin pigmentation: melanocytes modulate skin color in response to stress. Federation of American Societies for Experimental Biology Journal, 1: 976–994. doi.org/10.1096/fj.06-6649rev.
Deng, W., Tan, Y., Wang, X., Xi, D., He, Y., Yang, S., Mao, H. and Gao, S. (2009). Molecular cloning, sequence characteristics, and polymorphism analyses of the tyrosinase-related protein 2/DOPAchrome tautomerase gene of black-boned sheep (Ovis aries). Genome, 52: 1001-1011. doi.org/10.1139/G09-078.
Fan, W., Boston, B.A., Kesterson, R.A., Hruby, V.J. and Cone, R.D. (1997). Role of melanocortinergic neurons in feeding and the agouti obesity syndrome. Nature, 385: 165-168. doi.org/10.1038/385165a0.
Han, J. L., Min, Y., Guo, T. T., Yue, Y. J., Liu, J. B., Niu, C. E., Wang, C. F. and Yang, B. H. (2015). Molecular characterization of two candidate genes associated with coat color in Tibetan sheep (Ovis arise). Journal of Integrative Agriculture, 14: 1390-1397. doi.org/10.1016/S2095-3119(14)60928-X.
Hubbard, J. K., Uy, J. A. C., Hauber, M. E., Hoekstra, H. E. and Safran, R. J. (2010). Vertebrate pigmentation: from underlying genes to adaptive function, Trends Genetics, 26: 231–239. doi.org/10.1016/j.tig.2010.02.002.
Inagaki, K., Suzuki, T., Ito, S., Suzuki, N., Adachi, K., Okuyama, T., Nakata, Y., Shimizu, H., Matsuura, H. and Oono, T. 2006. Oculocutaneous albinism type 4: six novel mutations in the membrane‐associated transporter protein gene and their phenotypes. Pigment Cell Research, 19: 451-453. doi.org/10.1111/j.1600-0749.2006.00332.x.
Ko, J. M., Yang, J. A., Jeong, S. Y. and Kim, H. J. (2012). Mutation spectrum of the TYR and SLC45A2 genes in patients with oculocutaneous albinism. Molecular Medicine Reports, 5: 943-948. doi.org/10.3892/mmr.2012.764.
Koseniuk, A., Ropka-Molik, K., Rubiś, D. and Smołucha, G. (2018). Genetic background of coat colour in sheep. Archives Animal Breeding, 61(2): 173-178. doi.org/10.5194/aab-61-173-2018.
Lamason, R. L., Mohideen,M. A. P., Mest, J. R., Wong, A. C., Norton, H. L., Aros, M. C., Jurynec, M. J., Mao, X., Humphreville, V. R. and Humbert, J. E. (2005). SLC24A5, a putative cation exchanger, affects pigmentation in zebrafish and humans. Science, 310: 1782-1786. doi.org/10.1126/science.1116238.
Li, H. T., Zeng, X. C., Zhang, W. X., Zhao, H. X., Hui, W. Q., Liu, G., Luo, Y., Ban, Q., Zhao, Z. S. and Jia, B. (2013). Analysis of the relationship between polymorphism/mRNA expression levels of MC1R and ASIP and coat color phenotype in Kazakh sheep. Chinese Journal of Animal and Veterinary Sciences, 44(3): 366-75. https://www.xmsyxb.com/EN/Y2013/V44/I3/366
Moradi, K.,, Mustafa Muhaghegh Dolatabady, M.and Habibizad, J. (2024). Expression analysis of candidate genes in white and black spots of skin tissue in Lori Bakhtiari sheep. Iranian Journal of Animal Science researh. (In press). doi.org/10.22067/ijasr.2023.83699.1166
Nan, H., Kraft, P., Hunter, D.J. and Han, J. (2009). Genetic variants in pigmentation genes, pigmentary phenotypes, and risk of skin cancer in Caucasians. International Journal of Cancer, 125: 909-917. doi.org/10.1002/ijc.24327.
Nazari-Ghadikolaei, A., Mehrabani-Yeganeh, H., Miarei-Aashtiani, S. R., Staiger, E. A., Rashidi, A., Huson, H. J. (2018). Genome-wide association studies identify candidate genes for coat color and mohair traits in the Iranian Markhoz goat. Frontiers in Genetics, 9: 105. doi.org/10.3389/fgene.2018.00105.
Newton, J., Cohen-Barak, O., Hagiwara, N., Gardner, J.M., Davisson, M.T., King, R.A. and Brilliant, M.H. (2001). Mutations in the human orthologue of the mouse underwhite gene (uw) underlie a new form of oculocutaneous albinism, OCA4. The American Journal of Human Genetics, 69: 981-988. doi.org/10.1086/324340
Norton, H. L., Kittles, R. A., Parra, E., McKeigue, P., Mao, X., Cheng, K., Canfield, V. A., Bradley, D. G., McEvoy, B. and Shriver, M. D. (2007). Genetic evidence for the convergent evolution of light skin in Europeans and East Asians. Molecular Biology and Evolution, 24: 710-722. doi.org/10.1093/molbev/msl203.
Parichy, D. M., Reedy, M. V. and Erickson, C. A. (2006). Regulation of melanoblast migration and differentation. The Pigmentary System and its disorders, 2nd Edn., Oxford University Press, Oxford,108–127. doi.org/10.1002/9780470987100.ch5.
Peñagaricano, F., Zorrilla, P., Naya, H., Robello, C. and Urioste, J. I. (2012). Gene expression analysis identifies new candidate genes associated with the development of black skin spots in Corriedale sheep. Journal of Applied Genetics, 53: 99-10. doi.org/10.1007/s13353-011-0066-9.
Peng, Y., Wang, Y., Wang, R., Geng, L., Ma, R., Zhang, C., Liu, Z., Gong, Y., Li, J. and Li, X. (2018). Exploring differentially expressed genes associated with coat color in goat skin using RNA-seq. Canadian Journal of Animal Science, 99: 357-366. doi.org/10.1139/cjas-2018-0026.
Pfaffl, M.W., Horgan, G.W. and Dempfle, L. (2002). Relative expression software tool (REST©) for group-wise comparison and statistical analysis of relative expression results in real-time PCR. Nucleic Acids Research, 30: e36-e36. doi.org/ 10.1093/nar/30.9.e36.
Renieri, C., Valbonesi, A., La Manna, V., Antonini, M., and Lauvergne, J. J. (2008). Inheritance of coat colour in Merino sheep, Small Ruminant Research, 74: 23–29. doi.org/10.1016/j.smallrumres.2007.03.001.
Robbins, L. S., Nadeau, J. H., Johnson, K. R., Kelly, M. A., Roselli- Rehfuss, L., Baack, E., Mountjoy, K. G., and Cone, R. D. (1993). Pigmentation phenotypes of variant extension locus alleles result from point mutations that alter MSH receptor function. Cell, 72: 827–834, 1993. doi.org/10.1016/0092-8674(93)90572-8.
Royo, L.J., Alvarez, I., Arranz, J., Fernández, I., Rodríguez, A., Pérez‐Pardal, L. and Goyache, F. (2008). Differences in the expression of the ASIP gene are involved in the recessive black coat colour pattern in sheep: evidence from the rare Xalda sheep breed. Animal Genetics, 39: 290-293. doi.org/10.1111/j.1365-2052.2008.01712.x.
Saadat Nouri, M. and Siah Mansour, P. (1989). Principles of maintenance and raising sheep. Fourth edition, Tehran, Farhang Publications. (In Persian).
Saternus, R., Pilz, S., Gräber, S., Kleber, M., März, W., Vogt, T. and Reichrath, J. (2015). A closer look at evolution: Variants (SNPs) of genes involved in skin pigmentation, including EXOC2, TYR, TYRP1, and DCT ,are associated with 25 (OH) D serum concentration. Endocrinology, 156: 39-47. doi.org/10.1210/en.2014-1238.
Searle, A. G. (1968). Comparative genetics of coat colour in mammals. Comparative genetics of coat colour in mammals. London: Logos Press Ltd. in association with Elek Books Ltd. Distributed by Academic Press, New Tork and London.
Solano, F. (2014). Melanins: skin pigments and much more—types, structural models, biological functions, and formation routes. New Journal of Science, 1-28. doi.org/10.1155/2014/498276.
Song, X., Xu, C., Liu, Z., Yue, Z., Liu, L., Yang, T., Cong, B. and Yang, F. (2017). Comparative transcriptome analysis of mink (Neovison vison) skin reveals the key genes involved in the melanogenesis of black and white coat colour. Scientific Reports, 7: 1-11. doi.org/10.1038/s41598-017-12754-0.
Thornton, B. and Basu, C. (2011). Real‐time PCR (qPCR) primer design using free online software. Biochemistry and Molecular Biology Education, 39(2): 145-154. https://doi.org/10.1002/bmb.20461.
Vasu, M., Ahlawat, S., Chhabra, P., Sharma, U., Arora, R., Sharma, R., Mir, M. A. and Singh, M. K. (2024). Genetic insights into fiber quality, coat color and adaptation in Changthangi and Muzzafarnagri sheep: A comparative skin transcriptome analysis. Gene, 891:147826. doi.org/10.1016/j.gene.2023.147826.
Wang, H., Xue, L., Li, Y., Zhao, B., Chen, T., Liu, Y., Chang, L. and Wang, J. (2016). Distribution and expression of SLC45A2 in the skin of sheep with different coat colors. Folia Histochemica et Cytobiologica, 54: 143-150. doi.org/10.5603/FHC.a2016.0015
Yamaguchi, Y., Brenner, M. and Hearing, V. J. (2007). The regulation of skin pigmentation. Journal of biological chemistry, 282: 27557-27561. doi.org/10.1074/jbc.R700026200.
Yao, L., Bao, A., Hong, W., Hou, C., Zhang, Z., Liang, X. and Aniwashi, J. (2019). Transcriptome profiling analysis reveals key genes of different coat color in sheep skin. Peer Journal, 7: e8077. doi.org/ 10.7717/peerj.8077.
Zhang, W., Jin, M., Lu, Z., Li, T., Wang, H., Yuan, Z. and Wei, C. (2023). Whole genome resequencing reveals selection signals related to wool color in sheep. Animals, 13(20): 3265. doi.org/10.3390/ani13203265.